Cyperaceae taxon details
Tricostularia exsul (C.B.Clarke) K.L.Wilson & R.L.Barrett
Lepidosperma exsul C.B.Clarke · unaccepted
Tricostularia neesii var. elatior Benth. · unaccepted
recent only
Not documented
Lectotype (of Lepidosperma exsul C.B.Clarke) G...
Lectotype (of Lepidosperma exsul C.B.Clarke) G 00195302, geounit Western Australia [details]
Description Perennial tufted herb, 0.4–1(–1.3) m high; plants clonal, forming dense tussocks 0.2–1.2 m across; rhizome thin,...
Description Perennial tufted herb, 0.4–1(–1.3) m high; plants clonal, forming dense tussocks 0.2–1.2 m across; rhizome thin, woody, short, branched, 1.6–2.3 mm diam., non- or slightly pseudobulbous; old leaf sheaths covering the rhizome, breaking apart into fine fibres with age, pale brown; roots sand-binding. Culms slender, rigid, erect, with 2 nodes, terete, finely striate, 35–90(–120) cm tall, 1.1–2.0 mm diam., smooth, glabrous, dull to bright medium green to yellow-green, base not enlarged, but noticeably thicker than apex. Leaves mostly basal, numerous, and 2 cauline; phyllotaxy loosely spirotristichous; sheath 33–80 mm long, 1.5–2.3 mm wide, broader than the leaf lamina, margins finely scabrid, otherwise glabrous, straw-coloured to reddish brown, dull, upper margin membranous, oblique, finely scabrid, ligule absent; cauline sheath tight around culm; pseudopetiole obscure or absent; basal leaves with a much reduced lamina, not exceeding the culms, lamina dorsiventral, 0.1–0.2 cm long, 0.4–0.6 mm wide; bases brown-membranous, not dividing; lamina ± linear or narrowly triangular, ± flat, ± flexuous, old leaf tips not curling, finely multi-striate, margins very finely scabrid (only visible at (40×), otherwise glabrous, brown, concolorous, weakly keeled along mid-nerve, margins not recurved, apex acute; lowest leaves reduced to a sheath, lacking a lamina, pale to reddish brown. Inflorescence elongate, panicle-like, narrow, somewhat flexuous, with 5–9 nodes, 8–12 cm long, 4–9 mm wide, sometimes interrupted at the base, axis green or yellow-green, not glaucous; bracts reduced leaf-like, ± equal the spikelet clusters, sheaths open, partly enveloping the higher bracts, gradually reducing along the inflorescence, basal bract much shorter than inflorescence, 20–30 mm long; branches erect, with usually 3 but up to 7 spikelets, spikelet(s) on short, compressed, scabrous peduncles 1–3 mm long, arising in each bract axil; basal branchlet 20–29 mm long; spikelet prophyll present, sheath 6.2–8.4 mm long, brown, with 3 distinct veins and many fine striations, partially enclosing the spikelet, with a slender lamina 3.3–4.6 mm long, brown. Spikelets brown, lanceolate, 8.5–11 mm long, 2.1–3.3 mm wide, subterete, with 5 or 6 glumes, only topmost 3 fertile, the lower flower male, sterile, the second flower male, fertile, the upper flower bisexual, fertile, sometimes with a highly reduced glume above the florets; glumes distichous, basal glume 3.6–4.2 mm long, fertile glumes 7.0–8.8 mm long, 1.3–2.2 mm wide, membranous, pale reddish brown, sometimes translucent, sparsely scabrous on the keel and margins, lamina glabrous, lanceolate, acuminate, with a mucro 0.3–1.0 mm long, keel distinct; rachilla compact and not sinuous in fruiting spikelets. Perianth segments 6, whitish, minute, 0.7–2.0 mm long, ± filiform, expanded below, tapering to a fine point, sparsely scabrous. Stamens 3; anther connective 5.9–7.8 mm long, subulate; anthers yellow, 3.6–4.1 mm long, with a glabrous apical appendage 0.8–1.2 mm long. Style trifid, base 3.6–4.1 mm long, glabrous, slender above and below (dilated on nutlet apex), branches 2.8–3.2 mm long, hispidulous. Nutlet shortly stipitate, stipe 0.5–0.6 mm long, somewhat constricted, broad ellipsoid, obscurely trigonous, dull, mid-brown to dark brown, 2.5–2.8 mm long including the stipe and style base, 1.3–1.5 mm diam., faces smooth, puberulous, densely so near apex, epidermal cells square–hexagonal, inconspicuous, with three fine white ribs, style base distinctly enlarged, thick, persistent, shortly conical; embryo not examined. Photosynthetic pathway not examined. [details]
Cyperaceae Working Group. (2025). [see How to cite]. Global Cyperaceae Database. Tricostularia exsul (C.B.Clarke) K.L.Wilson & R.L.Barrett. Accessed at: https://www.cyperaceae.org/aphia.php?p=taxdetails&id=1825562 on 2025-09-12
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Nomenclature
original description
(of Lepidosperma exsul C.B.Clarke) Clarke, C. B. (1908). New genera and species of Cyperaceae. <em>Bulletin of miscellaneous information. Additional Series.</em> 8: 1-196., available online at https://www.biodiversitylibrary.org/item/104847
page(s): 47 [details]
original description (of Tricostularia neesii var. elatior Benth.) Bentham, G. (1878). Flora australiensis: a description of the plants of the Australian territory. vol. 7, pp. 2-806. L. Reeve, Covent Garden, London., available online at https://www.biodiversitylibrary.org/page/11265831
page(s): 384 [details]
new combination reference Barrett, R. L.; Wilson, K. L. (2012). A review of the genus Lepidosperma Labill. (Cyperaceae: Schoeneae). Australian Systematic Botany, 25(4): 225-294., available online at https://doi.org/10.1071/SB11037
page(s): 286 [details]
page(s): 47 [details]
original description (of Tricostularia neesii var. elatior Benth.) Bentham, G. (1878). Flora australiensis: a description of the plants of the Australian territory. vol. 7, pp. 2-806. L. Reeve, Covent Garden, London., available online at https://www.biodiversitylibrary.org/page/11265831
page(s): 384 [details]
new combination reference Barrett, R. L.; Wilson, K. L. (2012). A review of the genus Lepidosperma Labill. (Cyperaceae: Schoeneae). Australian Systematic Botany, 25(4): 225-294., available online at https://doi.org/10.1071/SB11037
page(s): 286 [details]
Present Inaccurate Introduced: alien Containing type locality
Lectotype (of Lepidosperma exsul C.B.Clarke) G 00195302, geounit Western Australia [details]
From editor or global species database
Description Perennial tufted herb, 0.4–1(–1.3) m high; plants clonal, forming dense tussocks 0.2–1.2 m across; rhizome thin, woody, short, branched, 1.6–2.3 mm diam., non- or slightly pseudobulbous; old leaf sheaths covering the rhizome, breaking apart into fine fibres with age, pale brown; roots sand-binding. Culms slender, rigid, erect, with 2 nodes, terete, finely striate, 35–90(–120) cm tall, 1.1–2.0 mm diam., smooth, glabrous, dull to bright medium green to yellow-green, base not enlarged, but noticeably thicker than apex. Leaves mostly basal, numerous, and 2 cauline; phyllotaxy loosely spirotristichous; sheath 33–80 mm long, 1.5–2.3 mm wide, broader than the leaf lamina, margins finely scabrid, otherwise glabrous, straw-coloured to reddish brown, dull, upper margin membranous, oblique, finely scabrid, ligule absent; cauline sheath tight around culm; pseudopetiole obscure or absent; basal leaves with a much reduced lamina, not exceeding the culms, lamina dorsiventral, 0.1–0.2 cm long, 0.4–0.6 mm wide; bases brown-membranous, not dividing; lamina ± linear or narrowly triangular, ± flat, ± flexuous, old leaf tips not curling, finely multi-striate, margins very finely scabrid (only visible at (40×), otherwise glabrous, brown, concolorous, weakly keeled along mid-nerve, margins not recurved, apex acute; lowest leaves reduced to a sheath, lacking a lamina, pale to reddish brown. Inflorescence elongate, panicle-like, narrow, somewhat flexuous, with 5–9 nodes, 8–12 cm long, 4–9 mm wide, sometimes interrupted at the base, axis green or yellow-green, not glaucous; bracts reduced leaf-like, ± equal the spikelet clusters, sheaths open, partly enveloping the higher bracts, gradually reducing along the inflorescence, basal bract much shorter than inflorescence, 20–30 mm long; branches erect, with usually 3 but up to 7 spikelets, spikelet(s) on short, compressed, scabrous peduncles 1–3 mm long, arising in each bract axil; basal branchlet 20–29 mm long; spikelet prophyll present, sheath 6.2–8.4 mm long, brown, with 3 distinct veins and many fine striations, partially enclosing the spikelet, with a slender lamina 3.3–4.6 mm long, brown. Spikelets brown, lanceolate, 8.5–11 mm long, 2.1–3.3 mm wide, subterete, with 5 or 6 glumes, only topmost 3 fertile, the lower flower male, sterile, the second flower male, fertile, the upper flower bisexual, fertile, sometimes with a highly reduced glume above the florets; glumes distichous, basal glume 3.6–4.2 mm long, fertile glumes 7.0–8.8 mm long, 1.3–2.2 mm wide, membranous, pale reddish brown, sometimes translucent, sparsely scabrous on the keel and margins, lamina glabrous, lanceolate, acuminate, with a mucro 0.3–1.0 mm long, keel distinct; rachilla compact and not sinuous in fruiting spikelets. Perianth segments 6, whitish, minute, 0.7–2.0 mm long, ± filiform, expanded below, tapering to a fine point, sparsely scabrous. Stamens 3; anther connective 5.9–7.8 mm long, subulate; anthers yellow, 3.6–4.1 mm long, with a glabrous apical appendage 0.8–1.2 mm long. Style trifid, base 3.6–4.1 mm long, glabrous, slender above and below (dilated on nutlet apex), branches 2.8–3.2 mm long, hispidulous. Nutlet shortly stipitate, stipe 0.5–0.6 mm long, somewhat constricted, broad ellipsoid, obscurely trigonous, dull, mid-brown to dark brown, 2.5–2.8 mm long including the stipe and style base, 1.3–1.5 mm diam., faces smooth, puberulous, densely so near apex, epidermal cells square–hexagonal, inconspicuous, with three fine white ribs, style base distinctly enlarged, thick, persistent, shortly conical; embryo not examined. Photosynthetic pathway not examined. [details]
To Biodiversity Heritage Library (2 publications) (from synonym Lepidosperma exsul C.B.Clarke)
To GenBank (1 nucleotides; 0 proteins) (from synonym Lepidosperma exsul C.B.Clarke)
To Genève Herbarium (Lepidosperma exsul G00195301 isotype 1) (from synonym Lepidosperma exsul C.B.Clarke)
To Genève Herbarium (Lepidosperma exsul G00195302 holotype 1) (from synonym Lepidosperma exsul C.B.Clarke)
To Genève Herbarium (Lepidosperma exsul G00195303 isotype 1) (from synonym Lepidosperma exsul C.B.Clarke)
To International Plant Names Index (IPNI) (from synonym Lepidosperma exsul C.B.Clarke)
To International Plant Names Index (IPNI) (from synonym Tricostularia neesii var. elatior Benth.)
To Plants of the World Online (from synonym Lepidosperma exsul C.B.Clarke)
To Plants of the World Online (from synonym Tricostularia neesii var. elatior Benth.)
To Royal Botanic Gardens, Kew (Tricostularia neesii var. elatior K000960117 type 1) (from synonym Tricostularia neesii var. elatior Benth.)
To GenBank (1 nucleotides; 0 proteins) (from synonym Lepidosperma exsul C.B.Clarke)
To Genève Herbarium (Lepidosperma exsul G00195301 isotype 1) (from synonym Lepidosperma exsul C.B.Clarke)
To Genève Herbarium (Lepidosperma exsul G00195302 holotype 1) (from synonym Lepidosperma exsul C.B.Clarke)
To Genève Herbarium (Lepidosperma exsul G00195303 isotype 1) (from synonym Lepidosperma exsul C.B.Clarke)
To International Plant Names Index (IPNI) (from synonym Lepidosperma exsul C.B.Clarke)
To International Plant Names Index (IPNI) (from synonym Tricostularia neesii var. elatior Benth.)
To Plants of the World Online (from synonym Lepidosperma exsul C.B.Clarke)
To Plants of the World Online (from synonym Tricostularia neesii var. elatior Benth.)
To Royal Botanic Gardens, Kew (Tricostularia neesii var. elatior K000960117 type 1) (from synonym Tricostularia neesii var. elatior Benth.)
