Cyperaceae taxon details
Carex complexa Reznicek & S.González
1676690 (urn:lsid:marinespecies.org:taxname:1676690)
accepted
Species
terrestrial
Reznicek, A. A.; González‐Elizondo, M. D. S.; Hahn, M.; Garner, M.; Hipp, A. L. (2021). Monograph of <i>Carex</i> section <i>Schiedeanae</i> (Cyperaceae): Unexpected taxonomic and ecological diversity in a Mexican sedge clade. <em>Journal of Systematics and Evolution.</em> 59(4): 698-725., available online at https://doi.org/10.1111/jse.12741
page(s): 712 [details]
page(s): 712 [details]
Holotype MICH, geounit Mexico
Holotype MICH, geounit Mexico [details]
Description Habit densely cespitose in large clumps from very short, stout, fibrillose rhizomes. Culms 20–60 cm tall, erect,...
Description Habit densely cespitose in large clumps from very short, stout, fibrillose rhizomes. Culms 20–60 cm tall, erect, ±equaling to longer than the leaves, trigonous, finely papillose and somewhat antrorsely scabrous-angled distally; bladeless basal sheaths promptly fibrillous and dark-colored (as well as the leaves of previous years). Leaves 6–8 (–10), clustered on the proximal 1/4–1/10 of the culms; blades coriaceous, 15–40 cm long, (1.2–) 1.5–3 mm wide, with strongly revolute margins, attenuate, becoming ±channeled distally and finally triangular and curled at the apex, minutely papillose adaxially and usually also abaxially, the midrib prominent on the abaxial side, the margins and midrib antrorsely scabrous distally, light-green to yellow green; leaf sheaths tight, minutely papillose, pale green to stramineous dorsally, whitish or yellowish to reddish-brown stained, ±hyaline ventrally, truncate to slightly concave at the ±thickened apex; ligules truncate or retuse, reduced to the free portion of ca. 0.2 mm wide, white to yellowish. Vegetative shoots abundant in the clumps, 2–6 cm tall with 6–10 leaves. Inflorescences 1.5–6 cm long, 0.6–1 cm wide, oblong, with 5–11 branches, the distal 2–6 branches approximate, sessile, and bearing simple androgynous spikes, the proximal 1–5 branches compound (or simple spikes in depauperate inflorescences), the proximalmost usually a 2–8-spiked panicle, proximalmost inflorescence branch 3.5–18.5 mm distant, subsessile or short (up to 13.5 mm) peduncled; proximalmost bracts 3.7–15.5 cm long, foliaceous, revolute, antrorsely scabrous-margined to the base, the distal bracts gradually reduced; terminal spike 5–9 mm long, pistillate portion 2–4.5 mm long, 3.1–4.3 mm wide, 4–8 flowered, staminate portion 2.6–5.9 mm long, 0.7–1 mm wide, 10–24-flowered, often relatively inconspicuous, extending 0–3.5 mm beyond the perigynia; cladoprophylls ±tubular, greenish, and finely pubescent on the primary branches, more distended and pubescent on the secondary branches. pistillate scales 1.5–2.8 (–3.5) mm long, 0.8-1.3 mm wide, all but the proximalmost much shorter than the perigynia, ovate to ovate-lanceolate, glabrous, white to reddish-brown-tinged with a 1–3-nerved green center and hyaline margins, acute to acuminate, the proximalmost occasionally with an awn up to 2.2 mm. Staminate scales 1.4–1.8 mm long, ca. 0.9–1 mm wide, ovate, obtuse to acute, ±cucullate, whitish-hyaline or somewhat reddish-brown-tinged with a 1–3-nerved green center. Perigynia 2.4–3 mm long, 0.7–1.1 mm wide, ascending to spreading-ascending, trigonous with elliptic sides, adaxial side ±concave, nerveless or inconspicuously nerved, abaxial sides flat, 5–8-nerved, tapering to the base, green, herbaceous, papillose, not tuberculate, hairs appressed-retrorse with some ±spreading, 0.04–0.07 mm long, distributed over the entire surface, strongly flattened, narrowly deltate to deltate, whitish; tapered into a beak; beaks (0.5–) 0.6–1.1 mm long, ca. 0.3-0.4 mm wide, widely green margined, with false dorsal suture conspicuous, ±deeply bidentate at the apex with teeth 0.2–0.6 mm long. Achenes 1.7–2 mm long, 0.7–1.1 mm wide, trigonous with slightly concave, elliptic sides, pale-brown, finely papillose, sessile, apiculus essentially lacking; style straight, glabrous, stigmas ca. 0.3–0.7 mm long, coiled, finely papillose to ±smooth (papillae shorter than the width of the stigmas). Anthers 0.5–0.7 mm long, filaments short-exserted to 0.1–0.7 mm, flattened, ca. 0.1–0.3 mm wide, about ½ as wide as the anther, stiffly curved, yellow,
darkening apically after the anthers are shed. [details]
darkening apically after the anthers are shed. [details]
Cyperaceae Working Group. (2025). [see How to cite]. Global Cyperaceae Database. Carex complexa Reznicek & S.González. Accessed at: https://cyperaceae.org/aphia.php?p=taxdetails&id=1676690 on 2025-09-12
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2024-12-10 11:47:54Z
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Nomenclature
original description
Reznicek, A. A.; González‐Elizondo, M. D. S.; Hahn, M.; Garner, M.; Hipp, A. L. (2021). Monograph of <i>Carex</i> section <i>Schiedeanae</i> (Cyperaceae): Unexpected taxonomic and ecological diversity in a Mexican sedge clade. <em>Journal of Systematics and Evolution.</em> 59(4): 698-725., available online at https://doi.org/10.1111/jse.12741
page(s): 712 [details]
basis of record Plants of the World Online (POWO). , available online at https://powo.science.kew.org/ [details]
page(s): 712 [details]
basis of record Plants of the World Online (POWO). , available online at https://powo.science.kew.org/ [details]
Present Inaccurate Introduced: alien Containing type locality
Holotype MICH, geounit Mexico [details]
From editor or global species database
Description Habit densely cespitose in large clumps from very short, stout, fibrillose rhizomes. Culms 20–60 cm tall, erect, ±equaling to longer than the leaves, trigonous, finely papillose and somewhat antrorsely scabrous-angled distally; bladeless basal sheaths promptly fibrillous and dark-colored (as well as the leaves of previous years). Leaves 6–8 (–10), clustered on the proximal 1/4–1/10 of the culms; blades coriaceous, 15–40 cm long, (1.2–) 1.5–3 mm wide, with strongly revolute margins, attenuate, becoming ±channeled distally and finally triangular and curled at the apex, minutely papillose adaxially and usually also abaxially, the midrib prominent on the abaxial side, the margins and midrib antrorsely scabrous distally, light-green to yellow green; leaf sheaths tight, minutely papillose, pale green to stramineous dorsally, whitish or yellowish to reddish-brown stained, ±hyaline ventrally, truncate to slightly concave at the ±thickened apex; ligules truncate or retuse, reduced to the free portion of ca. 0.2 mm wide, white to yellowish. Vegetative shoots abundant in the clumps, 2–6 cm tall with 6–10 leaves. Inflorescences 1.5–6 cm long, 0.6–1 cm wide, oblong, with 5–11 branches, the distal 2–6 branches approximate, sessile, and bearing simple androgynous spikes, the proximal 1–5 branches compound (or simple spikes in depauperate inflorescences), the proximalmost usually a 2–8-spiked panicle, proximalmost inflorescence branch 3.5–18.5 mm distant, subsessile or short (up to 13.5 mm) peduncled; proximalmost bracts 3.7–15.5 cm long, foliaceous, revolute, antrorsely scabrous-margined to the base, the distal bracts gradually reduced; terminal spike 5–9 mm long, pistillate portion 2–4.5 mm long, 3.1–4.3 mm wide, 4–8 flowered, staminate portion 2.6–5.9 mm long, 0.7–1 mm wide, 10–24-flowered, often relatively inconspicuous, extending 0–3.5 mm beyond the perigynia; cladoprophylls ±tubular, greenish, and finely pubescent on the primary branches, more distended and pubescent on the secondary branches. pistillate scales 1.5–2.8 (–3.5) mm long, 0.8-1.3 mm wide, all but the proximalmost much shorter than the perigynia, ovate to ovate-lanceolate, glabrous, white to reddish-brown-tinged with a 1–3-nerved green center and hyaline margins, acute to acuminate, the proximalmost occasionally with an awn up to 2.2 mm. Staminate scales 1.4–1.8 mm long, ca. 0.9–1 mm wide, ovate, obtuse to acute, ±cucullate, whitish-hyaline or somewhat reddish-brown-tinged with a 1–3-nerved green center. Perigynia 2.4–3 mm long, 0.7–1.1 mm wide, ascending to spreading-ascending, trigonous with elliptic sides, adaxial side ±concave, nerveless or inconspicuously nerved, abaxial sides flat, 5–8-nerved, tapering to the base, green, herbaceous, papillose, not tuberculate, hairs appressed-retrorse with some ±spreading, 0.04–0.07 mm long, distributed over the entire surface, strongly flattened, narrowly deltate to deltate, whitish; tapered into a beak; beaks (0.5–) 0.6–1.1 mm long, ca. 0.3-0.4 mm wide, widely green margined, with false dorsal suture conspicuous, ±deeply bidentate at the apex with teeth 0.2–0.6 mm long. Achenes 1.7–2 mm long, 0.7–1.1 mm wide, trigonous with slightly concave, elliptic sides, pale-brown, finely papillose, sessile, apiculus essentially lacking; style straight, glabrous, stigmas ca. 0.3–0.7 mm long, coiled, finely papillose to ±smooth (papillae shorter than the width of the stigmas). Anthers 0.5–0.7 mm long, filaments short-exserted to 0.1–0.7 mm, flattened, ca. 0.1–0.3 mm wide, about ½ as wide as the anther, stiffly curved, yellow, darkening apically after the anthers are shed. [details]
