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Cyperaceae taxon details

Carex pilulifera L.

1677973  (urn:lsid:marinespecies.org:taxname:1677973)

  1. Biota
  2. Plantae (Kingdom)
  3. Viridiplantae (Subkingdom)
  4. Streptophyta (Infrakingdom)
  5. Tracheophyta (Phylum (Division))
  6. Spermatophytina (Subphylum (Subdivision))
  7. Magnoliopsida (Class)
  8. Lilianae (Superorder)
  9. Poales (Order)
  10. Cyperaceae (Family)
  11. Cariceae (Tribe)
  12. Carex (Genus)
  13. Carex subg. Carex (Subgenus)
  14. Carex sect. Acrocystis (Section)
  15. Carex pilulifera (Species)
accepted
Species

Ordering

  • Alphabetically
  • By status

Children Display

  1. Subspecies Carex pilulifera subsp. azorica (J.Gay) Franco & Rocha Afonso
  2. Subspecies Carex pilulifera subsp. pilulifera
  3. Forma Carex pilulifera f. americana Boeckeler accepted as Carex albicans var. albicans
  4. Forma Carex pilulifera f. fallax Vatke ex Asch. & Graebn. accepted as Carex pilulifera subsp. pilulifera
  5. Forma Carex pilulifera f. fuscidula Waisb. accepted as Carex pilulifera subsp. pilulifera
  6. Forma Carex pilulifera f. golenzii Asch. & Graebn. accepted as Carex pilulifera subsp. pilulifera
  7. Forma Carex pilulifera f. laxa Waisb. accepted as Carex pilulifera subsp. pilulifera
  8. Forma Carex pilulifera f. longibracteata Lange accepted as Carex pilulifera subsp. pilulifera
  9. Forma Carex pilulifera f. major Wirtg. ex Herrenkohl accepted as Carex pilulifera subsp. pilulifera
  10. Forma Carex pilulifera f. pallida Peterm. accepted as Carex pilulifera subsp. pilulifera
  11. Forma Carex pilulifera f. pedunculata Waisb. accepted as Carex pilulifera subsp. pilulifera
  12. Forma Carex pilulifera f. refracta Waisb. accepted as Carex pilulifera subsp. pilulifera
  13. Variety Carex pilulifera var. azorica (J.Gay) Christ accepted as Carex pilulifera subsp. azorica (J.Gay) Franco & Rocha Afonso
  14. Variety Carex pilulifera var. bastardiana (DC.) Nyman accepted as Carex pilulifera subsp. pilulifera
  15. Variety Carex pilulifera var. bertolonii Pi.Savi ex Kük. accepted as Carex pilulifera subsp. pilulifera
  16. Variety Carex pilulifera var. bertolonii Pi.Savi accepted as Carex pilulifera L.
  17. Variety Carex pilulifera var. deflexa (Hornem.) Drejer accepted as Carex deflexa Hornem.
  18. Variety Carex pilulifera var. digyna Boeckeler accepted as Carex pilulifera subsp. pilulifera
  19. Variety Carex pilulifera var. hispidula Mérat accepted as Carex fimbriata Schkuhr
  20. Variety Carex pilulifera var. leesii Ridl. accepted as Carex pilulifera subsp. pilulifera
  21. Variety Carex pilulifera var. longibracteata Lange accepted as Carex pilulifera subsp. pilulifera
  22. Variety Carex pilulifera var. longibracteata Lange accepted as Carex pilulifera subsp. pilulifera
  23. Variety Carex pilulifera var. mucronata Peterm. accepted as Carex pilulifera subsp. pilulifera
  24. Variety Carex pilulifera var. novae-angliae Kurtz accepted as Carex rossii Boott
  25. Variety Carex pilulifera var. pallescens Beckh. accepted as Carex pilulifera subsp. pilulifera
  26. Variety Carex pilulifera var. vaginata Boeckeler accepted as Carex depressa subsp. transsilvanica (Schur) K.Richt. accepted as Carex transsilvanica Schur
  27. Variety Carex pilulifera var. vegeta Merino accepted as Carex pilulifera subsp. pilulifera
terrestrial
Linnaeus, C. (1753). Species plantarum, exhibentes plantas rite cognitas ad genera relatas cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas. Vol 2. Salvius, Stockholm., available online at https://www.biodiversitylibrary.org/page/358580
page(s): 976 [details] 
Description Rhizomes short, with densely tufted shoots, often decumbent; roots purple-brown; scales brown, soon becoming fibrous. Stems...  
Description Rhizomes short, with densely tufted shoots, often decumbent; roots purple-brown; scales brown, soon becoming fibrous. Stems 8–30 cm, ± rough above, wiry, sharply trigonous below spike, often arcuate and elongating in fruit. Leaves 5–20 cm × 1.5–2 mm, rough above, ± flat, papillose on upper surface, ± abruptly tapered to a short trigonous point, dull yellow-green or mid-green, pink-brown and persistent on decay, overwintering; hypostomous; sheaths pale red-brown or wine-red, becoming fibrous, inner face hyaline; apex straight; ligule 0.5–1 mm, rounded, with free margin minutely fimbriate. Inflorescence 1–4 cm; spikes clustered; bracts leaf-like or upper setaceous, rarely exceeding inflorescence, the lowest not sheathing. Male spike 1, (6–)8–15 mm; male glumes 3.5–4 mm, oblanceolate-elliptic, brown or chestnut, hyaline toward margin, with pale midrib; apex acute to acuminate. Female spikes 2–4, 4–8 mm, ovoid or subglobose, erect, sessile; female glumes 3–3.5 mm, broadly ovate, red-brown, hyaline toward margin, midrib green becoming yellow; apex acute. Utricles 2–3.5 mm, obovoid-ellipsoid, downy at least at apex, deep green; beak 0.3–0.5 mm, notched; stigmas 3; nut obovoid, trigonous. [details]

Description Densely caespitose. Stems 5-30(-40) cm, smooth or weakly (rarely strongly) scabrid above; basal sheaths yellowish- to...  
Description Densely caespitose. Stems 5-30(-40) cm, smooth or weakly (rarely strongly) scabrid above; basal sheaths yellowish- to reddish-brown, fibrous. Leaves 1 -5-3 mm wide, shorter than stems, flat or slightly canaliculate, pale, usually greyish-green. Male spike solitary, 5-15 x c. 2 mm. Female spikes 2-4, 5-8 mm, globose to ovoid, sessile, overlapping or the lowest remote; lowest bract usually exceeding spike, leaf-like, usually patent, not sheathing. Female glumes acute or acuminate, dark, usually reddish-brown, with narrow scarious margin. Utricles 1-5-3-5 mm, obovoid, greyish-green, sparsely to densely puberulent, more or less veinless, abruptly contracted into an emarginate, usually conical beak not more than 0-5 mm.  [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 2: 976. 1753 [1 May 1753] to Sp. Pl. : 976 (1753),...  
Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 2: 976. 1753 [1 May 1753] to Sp. Pl. : 976 (1753), information provided by Alan E. on email Jun. 07 2021 More details could be found in [details]
Cyperaceae Working Group. (2025). [see How to cite]. Global Cyperaceae Database. Carex pilulifera L.. Accessed at: https://cyperaceae.org/aphia.php?p=taxdetails&id=1677973 on 2026-05-30
Date
action
by
2023-09-06 07:28:28Z
created
Jiménez-Mejías, Pedro
2024-12-10 11:47:54Z
unchecked
db_admin
2025-03-11 11:19:03Z
changed
Márquez-Corro, José Ignacio

Creative Commons License The webpage text is licensed under a Creative Commons Attribution 4.0 License


Nomenclature

original description Linnaeus, C. (1753). Species plantarum, exhibentes plantas rite cognitas ad genera relatas cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas. Vol 2. Salvius, Stockholm., available online at https://www.biodiversitylibrary.org/page/358580
page(s): 976 [details] 

original description (of Carex triceps Schrank) Schrank, F. P. (1789). Baiersche Flora. , available online at https://bibdigital.rjb.csic.es/viewer/11265#page=1&viewer=picture&o=bookmark&n=0&q=
page(s): 287 [details] 

basis of record Plants of the World Online (POWO). , available online at https://powo.science.kew.org/ [details] 

Taxonomy

taxonomy source Jermy, A.C.; Simpson, D.A.; Foley, M.J.Y.; Porter, M.S. (2007). Sedges of the British Isles. 554 pp. Botanical Society of Britain & Ireland, United Kingdom. [details] Available for editors  PDF available [request]

taxonomy source Luceño, M., Castroviejo, S., Jiménez-Mejías, P. (2008). Cyperaceae. In: Castroviejo, Luceño, M., Galán, A., Jiménez-Mejías, P., Cabezas, F., Medina, L. (eds). Flora iberica. Plantas vasculares de la Península Ibérica e Islas Baleares. Vol. XVIII. Madrid: CSIC., available online at https://bibdigital.rjb.csic.es/records/item/12958-flora-iberica-vol-18-cyperaceae-pontederiaceae?offset=, http://www.floraiberica.es/PHP/familias_lista_.php?familia=Cyperaceae [details] 

 
 Present  Inaccurate  Introduced: alien  Containing type locality 
   

Syntype L 0052714, geounit Europe [details]
From editor or global species database
Description Rhizomes short, with densely tufted shoots, often decumbent; roots purple-brown; scales brown, soon becoming fibrous. Stems 8–30 cm, ± rough above, wiry, sharply trigonous below spike, often arcuate and elongating in fruit. Leaves 5–20 cm × 1.5–2 mm, rough above, ± flat, papillose on upper surface, ± abruptly tapered to a short trigonous point, dull yellow-green or mid-green, pink-brown and persistent on decay, overwintering; hypostomous; sheaths pale red-brown or wine-red, becoming fibrous, inner face hyaline; apex straight; ligule 0.5–1 mm, rounded, with free margin minutely fimbriate. Inflorescence 1–4 cm; spikes clustered; bracts leaf-like or upper setaceous, rarely exceeding inflorescence, the lowest not sheathing. Male spike 1, (6–)8–15 mm; male glumes 3.5–4 mm, oblanceolate-elliptic, brown or chestnut, hyaline toward margin, with pale midrib; apex acute to acuminate. Female spikes 2–4, 4–8 mm, ovoid or subglobose, erect, sessile; female glumes 3–3.5 mm, broadly ovate, red-brown, hyaline toward margin, midrib green becoming yellow; apex acute. Utricles 2–3.5 mm, obovoid-ellipsoid, downy at least at apex, deep green; beak 0.3–0.5 mm, notched; stigmas 3; nut obovoid, trigonous. [details]

Description Densely caespitose. Stems 5-30(-40) cm, smooth or weakly (rarely strongly) scabrid above; basal sheaths yellowish- to reddish-brown, fibrous. Leaves 1 -5-3 mm wide, shorter than stems, flat or slightly canaliculate, pale, usually greyish-green. Male spike solitary, 5-15 x c. 2 mm. Female spikes 2-4, 5-8 mm, globose to ovoid, sessile, overlapping or the lowest remote; lowest bract usually exceeding spike, leaf-like, usually patent, not sheathing. Female glumes acute or acuminate, dark, usually reddish-brown, with narrow scarious margin. Utricles 1-5-3-5 mm, obovoid, greyish-green, sparsely to densely puberulent, more or less veinless, abruptly contracted into an emarginate, usually conical beak not more than 0-5 mm.  [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 2: 976. 1753 [1 May 1753] to Sp. Pl. : 976 (1753), information provided by Alan E. on email Jun. 07 2021 More details could be found in [details]
    Definitions

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To Biodiversity Heritage Library (1 publication) (from synonym Trasus piluliferus (L.) Gray)
To Biodiversity Heritage Library (169 publications) (from synonym Carex triceps Schrank)
To Biodiversity Heritage Library (615 publications) 
To European Nucleotide Archive, ENA (Carex pilulifera) 
To International Plant Names Index (IPNI) (from synonym Carex triceps Schrank)
To International Plant Names Index (IPNI) 
To International Plant Names Index (IPNI) (from synonym Bitteria pilulifera (L.) Fedde & J.Schust.)
To International Plant Names Index (IPNI) (from synonym Trasus piluliferus (L.) Gray)
To Plants of the World Online (from synonym Trasus piluliferus (L.) Gray)
To Plants of the World Online 
To Plants of the World Online (from synonym Carex triceps Schrank)
To Plants of the World Online (from synonym Bitteria pilulifera (L.) Fedde & J.Schust.)
To The Plant List (from synonym Carex pilulifera var. bertolonii Pi.Savi)
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