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Cyperaceae taxon details

Carex lupulina Muhl. ex Willd.

1677569 (urn:lsid:marinespecies.org:taxname:1677569)

  1. Biota
  2. Plantae (Kingdom)
  3. Viridiplantae (Subkingdom)
  4. Streptophyta (Infrakingdom)
  5. Tracheophyta (Phylum (Division))
  6. Spermatophytina (Subphylum (Subdivision))
  7. Magnoliopsida (Class)
  8. Lilianae (Superorder)
  9. Poales (Order)
  10. Cyperaceae (Family)
  11. Cyperoideae (Subfamily)
  12. Cariceae (Tribe)
  13. Carex (Genus)
  14. Carex subg. Carex (Subgenus)
  15. Carex sect. HirtaClade (Section)
  16. Carex lupulina (Species)
accepted
Species
Carex sect. Lupulinae (J.Carey) Mack.
 Carex subsect. Lupulinae J.Carey
Carex sect. HirtaClade
Carex lupulina var. albomarginata Sherff · unaccepted > heterotypic synonym
Carex lupulina var. androgyna Dewey · unaccepted > heterotypic synonym
Carex lupulina var. longipedunculata Dudley · unaccepted > heterotypic synonym
Carex lupulina var. pedunculata Dewey · unaccepted > later homonym (non A.Gray ex L.C.Beck (1833))
Carex lupulina var. pedunculata A.Gray ex L.C.Beck · unaccepted > heterotypic synonym
Carex lurida L.H.Bailey · unaccepted > misapplication
Carex lurida var. divergens L.H.Bailey · unaccepted > misapplication

Ordering

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Children Display

  1. Variety Carex lupulina var. albomarginata Sherff accepted as Carex lupulina Muhl. ex Willd. (unaccepted > heterotypic synonym)
  2. Variety Carex lupulina var. androgyna Dewey accepted as Carex lupulina Muhl. ex Willd. (unaccepted > heterotypic synonym)
  3. Variety Carex lupulina var. bella-villa (Dewey) L.H.Bailey accepted as Carex lupulina Muhl. ex Willd.
  4. Variety Carex lupulina var. gigantea (Rudge) Britton accepted as Carex gigantea Rudge
  5. Variety Carex lupulina var. gigantoidea Dewey accepted as Carex retrorsa Schwein.
  6. Variety Carex lupulina var. longipedunculata Dudley accepted as Carex lupulina Muhl. ex Willd. (unaccepted > heterotypic synonym)
  7. Variety Carex lupulina var. lupulina accepted as Carex lupulina Muhl. ex Willd.
  8. Variety Carex lupulina var. pedunculata Dewey accepted as Carex lupulina Muhl. ex Willd. (unaccepted > later homonym, non A.Gray ex L.C.Beck (1833))
  9. Variety Carex lupulina var. pedunculata A.Gray ex L.C.Beck accepted as Carex lupulina Muhl. ex Willd. (unaccepted > heterotypic synonym)
  10. Variety Carex lupulina var. polystachya Schwein. & Torr. accepted as Carex lupuliformis Sartwell ex Dewey (unaccepted > superseded combination, Variety elevated to the rank of species, but the name Carex polystachya Sw. ex Wahlenb. (1803) was already used, necessitating the creation of new name (Carex lupuliformis).)
terrestrial
(1805). Sp. Pl., ed. 4 [Willdenow] 4(1), available online at https://www.biodiversitylibrary.org/page/670960#page/1/mode/1up
page(s): 266 [details] 
Description Plants loosely cespitose or not, short-rhizomatous. Culms solitary or not, erect, 20–100(–130) cm. Leaves 4–8; basal...  
Description Plants loosely cespitose or not, short-rhizomatous. Culms solitary or not, erect, 20–100(–130) cm. Leaves 4–8; basal sheaths reddish to brownish; sheath of distal leaf (1.3–)1.7–10(–25) cm; ligules triangular, 3.5–18 mm; blades 15–64 cm × 4–15 mm. Inflorescences 4–40 cm; peduncles of proximal spikes 0.5–20 cm, basal 2 peduncles 1–20 cm apart of terminal spike 0.5–6(–7) cm, shorter than or exceeding distal pistillate spike by 2 cm or less; bracts leafy, sheaths 0.5–15 cm, blades 13–55 cm × 3–11 mm. Spikes: proximal pistillate spikes (1–)2–5, distal spikes usually ± crowded, ascending, densely (4–)8–80-flowered, ovoid to cylindric, 1.5–6.5 × 1.3–3 cm; terminal staminate spikes 1(–2), 1.5–8.5 cm × 1–5 mm. Pistillate scales 1–7-veined, lanceolate to lanceolate-ovate, 6–15 × 1–2.7 mm, apex acute to awned, awns rough, to 6 mm. Anthers 3, 3–5.5 mm. Perigynia ascending, strongly 13–22-veined, ± stipitate, lance-ovoid, 11–19 × 3–6 mm, shiny, glabrous; beak conic, 6–10 mm. Achenes stipitate, rhomboid, flat to concave faces, angles smoothly thickened, not pointed or nobbed, 3–4(–4.5) × 1.7–2.6(–2.8) mm; style same texture as achene. [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 4: 266 1805 to Sp. Pl., ed. 4 , 4: 266 (1805), information...  
Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 4: 266 1805 to Sp. Pl., ed. 4 , 4: 266 (1805), information provided by Alan E. on email Jun. 07 2021 More details could be found in [details]
Cyperaceae Working Group. (2025). [see How to cite]. Global Cyperaceae Database. Carex lupulina Muhl. ex Willd.. Accessed at: https://cyperaceae.org/aphia.php?p=taxdetails&id=1677569 on 2026-09-29
Date
action
by
2023-09-06 07:28:28Z
created
Jiménez-Mejías, Pedro
2024-12-10 11:47:54Z
unchecked
db_admin
2025-03-11 11:19:03Z
changed
Márquez-Corro, José Ignacio

Creative Commons License The webpage text is licensed under a Creative Commons Attribution 4.0 License


Nomenclature

original description (1805). Sp. Pl., ed. 4 [Willdenow] 4(1), available online at https://www.biodiversitylibrary.org/page/670960#page/1/mode/1up
page(s): 266 [details] 

original description (of Carex lupulina var. pedunculata A.Gray ex L.C.Beck) Beck, Lewis C. (1833). Botany of the northern and middle states, or, A description of the plants found in the United States, north of Virginia arranged according to the natural system, with a synopsis of the genera according to the Linnaean system--a sketch of the rudiments of botany, and a glossary of terms. 471 p., available online at https://www.biodiversitylibrary.org/page/16936860
page(s): 438 [details] 

original description (of Carex bella-villa Dewey) Dewey, C. (1866). Caricography. <em>American Journal of Science and Arts, ser. 2.</em> 41: 226-230., available online at https://www.biodiversitylibrary.org/page/36447954
page(s): 229 [details] 

original description (of Carex canadensis Dewey) Dewey, C. (1866). Caricography. <em>American Journal of Science and Arts, ser. 2.</em> 41: 226-230., available online at https://www.biodiversitylibrary.org/page/36447954
page(s): 229 [details] 

original description (of Carex lupulina var. longipedunculata Dudley) Dudley, W. R. (1886). The Cayuga flora. Part I, A catalogue of the phaenogamia growing without cultivation in the Cayuga Lake basin. <em>Bulletin of the Cornell University (Science).</em> 2: 1-196., available online at https://www.biodiversitylibrary.org/item/121545
page(s): 119 [details] 

original description (of Carex lupulina var. albomarginata Sherff) Sherff, Earl E. (1911). A new variety of Carex lupulina. <em>Bulletin of the Torrey Botanical Club.</em> 38: 481-482, pl. 26., available online at https://www.biodiversitylibrary.org/page/12617678
page(s): 482 [details] 

original description (of Carex lupulina var. androgyna Dewey) Wood, Alphonso. (1870). The American botanist and florist: including lessons in the structure, life, and growth of plants; together with a simple analytical flora, descriptive of the native and cultivated plants growing in the Atlantic division of the American union. 445 p., available online at https://babel.hathitrust.org/cgi/pt?id=ncs1.ark:/13960/t8x92qm3f&seq=536
page(s): 376; note: Link: https://archive.org/details/americanbotanist00wooduoft/americanbotanist00wooduoft/page/376/mode/2up [details] 

original description (of Carex lupulina var. pedunculata Dewey) Wood, Alphonso. (1870). The American botanist and florist: including lessons in the structure, life, and growth of plants; together with a simple analytical flora, descriptive of the native and cultivated plants growing in the Atlantic division of the American union. 445 p., available online at https://babel.hathitrust.org/cgi/pt?id=ncs1.ark:/13960/t8x92qm3f&seq=536
page(s): 376 [details] 

basis of record Plants of the World Online (POWO). , available online at https://powo.science.kew.org/ [details] 

 
 Present  Inaccurate  Introduced: alien  Containing type locality 
   

Isotype (of Carex lupulina var. albomarginata Sherff) GH 01684662, geounit Michigan [details]
Isotype (of Carex lupulina var. albomarginata Sherff) GH 01684663, geounit Michigan [details]
Isotype (of Carex lupulina var. albomarginata Sherff) OS
452974, geounit Michigan [details]
Nontype P 00306531, geounit Ohio [details]
Syntype (of Carex lupulina var. pedunculata A.Gray ex L.C.Beck) GH 00221804, geounit New York [details]
From editor or global species database
Description Plants loosely cespitose or not, short-rhizomatous. Culms solitary or not, erect, 20–100(–130) cm. Leaves 4–8; basal sheaths reddish to brownish; sheath of distal leaf (1.3–)1.7–10(–25) cm; ligules triangular, 3.5–18 mm; blades 15–64 cm × 4–15 mm. Inflorescences 4–40 cm; peduncles of proximal spikes 0.5–20 cm, basal 2 peduncles 1–20 cm apart of terminal spike 0.5–6(–7) cm, shorter than or exceeding distal pistillate spike by 2 cm or less; bracts leafy, sheaths 0.5–15 cm, blades 13–55 cm × 3–11 mm. Spikes: proximal pistillate spikes (1–)2–5, distal spikes usually ± crowded, ascending, densely (4–)8–80-flowered, ovoid to cylindric, 1.5–6.5 × 1.3–3 cm; terminal staminate spikes 1(–2), 1.5–8.5 cm × 1–5 mm. Pistillate scales 1–7-veined, lanceolate to lanceolate-ovate, 6–15 × 1–2.7 mm, apex acute to awned, awns rough, to 6 mm. Anthers 3, 3–5.5 mm. Perigynia ascending, strongly 13–22-veined, ± stipitate, lance-ovoid, 11–19 × 3–6 mm, shiny, glabrous; beak conic, 6–10 mm. Achenes stipitate, rhomboid, flat to concave faces, angles smoothly thickened, not pointed or nobbed, 3–4(–4.5) × 1.7–2.6(–2.8) mm; style same texture as achene. [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 4: 266 1805 to Sp. Pl., ed. 4 , 4: 266 (1805), information provided by Alan E. on email Jun. 07 2021 More details could be found in [details]
    Definitions

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Original illustration in Schkuhr. 
Potential type in herb. Wood (NY 2288650) (from synonym Carex lupulina var. androgyna Dewey)
Syntype at GH (GH00221804) (from synonym Carex lupulina var. pedunculata A.Gray ex L.C.Beck)
To Biodiversity Heritage Library (1 publication) (from synonym Carex lupulina var. pedunculata Dewey)
To Biodiversity Heritage Library (20 publications) (from synonym Carex canadensis Dewey)
To Biodiversity Heritage Library (273 publications) (from synonym Carex lurida L.H.Bailey)
To Biodiversity Heritage Library (355 publications) 
To European Nucleotide Archive, ENA (Carex lupulina) 
To European Nucleotide Archive, ENA (Carex lurida) (from synonym Carex lurida L.H.Bailey)
To GenBank (25 nucleotides; 12 proteins) 
To International Plant Names Index (IPNI) 
To International Plant Names Index (IPNI) (from synonym Carex canadensis Dewey)
To International Plant Names Index (IPNI) (from synonym Carex lurida L.H.Bailey)
To International Plant Names Index (IPNI) (from synonym Carex bella-villa Dewey)
To International Plant Names Index (IPNI) (from synonym Carex lupulina var. androgyna Dewey)
To International Plant Names Index (IPNI) (from synonym Carex lurida var. divergens L.H.Bailey)
To International Plant Names Index (IPNI) (from synonym Carex lupulina var. bella-villa (Dewey) L.H.Bailey)
To International Plant Names Index (IPNI) (from synonym Carex lupulina var. longipedunculata Dudley)
To International Plant Names Index (IPNI) (from synonym Carex gigantea var. lupulina (Muhl. ex Willd.) Farw.)
To International Plant Names Index (IPNI) (from synonym Carex lupulina var. albomarginata Sherff)
To International Plant Names Index (IPNI) (from synonym Carex gigantea f. bella-villa (Dewey) Farw.)
To International Plant Names Index (IPNI) (from synonym Carex lupulina var. pedunculata Dewey)
To IUCN Red List (Least Concern) 
To Plants of the World Online (from synonym Carex bella-villa Dewey)
To Plants of the World Online 
To Plants of the World Online (from synonym Carex canadensis Dewey)
To Plants of the World Online (from synonym Carex lurida L.H.Bailey)
To Plants of the World Online (from synonym Carex lupulina var. bella-villa (Dewey) L.H.Bailey)
To Plants of the World Online (from synonym Carex lupulina var. androgyna Dewey)
To Plants of the World Online (from synonym Carex lupulina var. albomarginata Sherff)
To Plants of the World Online (from synonym Carex gigantea var. lupulina (Muhl. ex Willd.) Farw.)
To Plants of the World Online (from synonym Carex lupulina var. longipedunculata Dudley)
To Plants of the World Online (from synonym Carex lupulina var. pedunculata Dewey)
To Plants of the World Online (from synonym Carex lurida var. divergens L.H.Bailey)
To Plants of the World Online (from synonym Carex gigantea f. bella-villa (Dewey) Farw.)
To The Plant List (from synonym Carex lupulina var. lupulina)
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